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General Description: Allium aaseae onion is an early spring-flowering member of the lily family. It is perennial with an underground bulb that is usually buried at least a couple inches below ground level in mature individuals. Bulb coat reticulations may or may not be evident. The scape is round to slightly flattened, not winged. The two linear, channeled leaves are 1 - 4 mm wide, at least twice as long as the scape and typically lying on the ground when observed in the field early in the season. Its six similar-looking tepals are pink, often richly so, but fading to white, 6 - 9.5 mm long, and with entire to obscurely or strongly denticulate margins. Stamens are shorter than the tepals, the undehisced anthers and pollen are yellow.
Technical Description: Bulb ovoid, outer coats brownish, usually with obscure reticulations, the cells of which are transversely elongate and intricately contorted, the inner coats white to pink or reddish; scape (3) 5 - 11 (15) cm long, terete or slightly flattened, not winged; leaves two per scape, linear, channeled, 1 - 3 (4) mm wide, entire or the margins obscurely denticulate, 2 or more times longer than the scape, green (i.e., not withering) at anthesis, tending to be deciduous at maturity; bracts of the inflorescence 2 (3), ovate, obtuse to acuminate; umbel 5- to 25-flowered, pedicels shorter than or equal the perianth; perianth segments (6) 7–9.5 mm long, lanceolate to elliptic, entire to erose to obscurely or strongly denticulate with minute glandular teeth, erect, flaring at the tips, bright pink, fading with age or pressing, rarely white; stamens 1/2 - 2/3 as long as the perianth; anthers yellow, pollen yellow; ovary crestless or with three minute 2-lobed central processes, style included, stigma punctate, entire, capsule crestless (McNeal 1993).
Diagnostic Characteristics: There are several onion species occurring within and near the range of A. aaseae. Allium aaseae is most likely to be confused with A. simillimum, especially at mid-elevations in the Boise Foothills, where their distributions nearly overlap. Populations with purple-mottled anthers may actually be hybrids between the two species. The following key, adopted from McNeal's (1993) key to the onions of southwestern Idaho, can be used to distinguish the two:
Allium simillimum. Perianth segments white with green or reddish midveins, sometimes flushed with pink; anthers purple or mottled purple and white, pollen white or grayish; denticulations, particularly on the inner perianth segments obvious under a hand lens and regularly distributed on the distal 2/3 of the segment; occurring above 4200 feet elevation on various substrates.
Allium aaseae. Perianth segments bright pink with rarely a white individual in an otherwise pink population; anthers yellow, pollen yellow, denticulations irregular in number and distribution on perianth segments, often missing on plants from Rebecca Sand Hills RNA; usually restricted to lacustrine sands of the Glenns Ferry Formation, generally below 3700 ft., except in Cartwright Canyon where occurring up to 5100 ft. elevation.
Infraspecific Taxa: There are no infraspecific taxa recognized for Allium aaseae.
Identification of this Taxon in Idaho: The bright pink flowers, yellow anthers, and restriction to course sandy substrates, generally below 3700 feet elevation, will distinguish Allium aaseae from the congener it is most likely to be confused, A. simillimum (dwarf onion). Additionally, the linear leaves of A. aaseae are at least twice as long as the scape, and typically lying on the ground when observed early in the spring. On well-developed plants, the leaves are also often much more succulent-looking than found in A. simillimum.
Hybridization involving A. aaseae has been suspected at least since the late 1970's. Recent research indicates hybridization is likely occurring between A. aaseae and A. simillimum at some populations. A recent taxonomic study by McNeal (1993) has determined Allium aaseae to be a distinct and valid species from A. simillimum. A genetic-based study by Smith (1995) substantiates McNeal's conclusion.
Global: Allium aaseae is endemic to southwestern Idaho. Population sizes vary from less than 100 to more than 35,000 plants. A rangewide population estimate for Allium aaseae is 400,000 plants. This is a conservative number for a number of reasons: 1) the full extent of a number of occurrences is unknown, and it is very likely additional plants occur in unsurveyed suitable habitat; 2) no population estimates are available for five occurrences and their contributions remain uncounted in the above rangewide tally; 3) for populations or subpopulations estimated at 10,000+ plants, only 10,000 were added to the tally; 4) although the majority of areas likely to support plants have been searched, some places, especially on private lands remain unsurveyed; 5) plants that are not flowering are difficult to see and their numbers are likely underestimated during field investigations. This conservative estimate of 400,000 plants reflects increased survey work for this species over the years. For instance, in 1978, Holsinger (1978) estimated less than 15,000 individuals rangewide, while ten years later Moseley and Caicco (1989) estimated 260,000 plants for 57 location sites. For the 68 occurrences of Allium aaseae, 17 (26%) are estimated to contain more than 10,000 individuals, 33 (49%) are estimated between 1000 - 10,000 plants, 12 (17%) are estimated between 100 - 1000 plants, and only one (1%) at fewer than 100. Abundance information is unknown in five (8%) instances.
Until the U. S. Fish and Wildlife Service recently revised their candidate system, Allium aaseae was a federal category 1 candidate for listing under the Endangered Species Act. Under the revised system, it is no longer a federal candidate species. Allium aaseae is a BLM Sensitive Species for Idaho. NatureServe, the parent organization for Natural Heritage Programs and Conservation Data Centers, ranks Allium aaseae as G3, a rank that includes taxa that are globally rare or uncommon, but not imperiled. Because Allium aaseae is endemic to Idaho, the state rank (S3) equals the global rank (G3). The Idaho Native Plant Society includes Allium aaseae on its list of globally rare plant taxa with a priority of 11, indicating threats are of low magnitude and non-imminent.
Substantial reduction in habitat and populations has occurred due to past land uses, although the exact amount is unknown. The main factors contributing to the serious conservation concern for Allium aaseae are its limited distribution, its restricted habitat requirements, and its location adjacent to a major population center making it subject to numerous threats (Moseley and Caicco 1989). Across its range, the sandy foothill habitats supporting Allium aaseae have been subject to four main land uses since European settlement: urban/suburbanization; livestock grazing; sand mining; and recreational uses, such as ORV's (off-road vehicle), equestrians, hikers, and mountain bikers. As a consequence of urban development in the Boise Foothills, portions of at least seven (11%) occurrences (007, 008, 020, 021, 033, 063, 064) have been destroyed by housing development projects in recent years. Twenty-two (34%) additional occurrences (004, 010, 011, 012, 013, 014, 019, 022, 023, 029, 030, 032, 036, 042, 047, 050, 054, 060, 061, 062, 065, 066) are known to occur at least partly on private land and in locations potentially subject to development, although future plans for these parcels are unknown. Additional private properties may also be subject to various future development. Segments of populations at the Hidden Hollow landfill (026, 027, 056, 057) may be potentially threatened over the long-term due to landfill expansion. Portions of other populations have also been lost over the years to sand mining, landfill activities and other disturbances. This permanent habitat loss and fragmentation resulting from urban developments, especially in the Boise Foothills, has accelerated in recent years, and is expected to continue. It also represents the most serious and probably difficult to resolve threat facing Allium aaseae.
The effects of livestock grazing on Allium aaseae are mainly indirect, principally the ecological decline of foothills habitat. Invasion by weedy annuals is largely the result of past disturbances associated with intensive livestock grazing and increases in fire frequency patterns. Livestock grazing is expected to remain a widespread land use throughout the range of Allium aaseae. Due to its early phenology, small and low growth habit, and typical occurrence in microsites with sparse associated forage, Allium aaseae is less prone to direct livestock impacts than many other native plants. Livestock use is not expected to result in the direct loss of onion habitat. However, in response to indirect effects such as fostering the invasion of cheatgrass (Bromus tectorum), storksbill (Erodium cicutarium), medusahead wildrye (Taeniatherum caput-medusae), rush skeleton weed (Chondrilla juncea) and other weeds, habitat quality will likely remain in decline for a long time in many places.
Management of the BLM's five ACEC's is designed to minimize impacts from livestock in these designated areas. Occurring mainly in the Pearl Mining District near Emmett, sand mining has destroyed or fragmented segments of several populations. Recreational impacts are generally local, although they may be more extensive where more concentrated or destructive. Portions of one occurrence (022) in Hulls Gulch has been destroyed by 4-wheel drive roads and associated destructive land rehabilitation actions. Portions of the large Sand Hollow population (034) are within the Little Gem Motorcycle Club area and have been impacted by off-trail riding disturbances. Four-wheel drive roads and ORV use are known to threaten at least parts of several other populations (such as 003, 014, 021 and 065). There is more uncertainty regarding potential impacts in other places within or near 4-wheel drive road and ORV-use areas. Other recreational users potentially threaten local sections of populations in the Hulls Gulch (011, 014, 022, 060), Military Reserve Park (006, 025, 058, 059), Camelsback Reserve Park (009) and other places as well (021, 065). As the population of the Boise area continues to grow, so will recreational demands and conflicts in the foothills.
Public land managing agencies, notably the BLM, have taken several steps for the conservation of Allium aaseae. Portions of five populations are located within BLM ACEC's (Cartwright Canyon, Sand-capped Knob, Sand Hollow, Willow Creek, and Woods Gulch), which were designated in 1993, primarily to protect Aase's onion. In addition, a disjunct population of Aase's onion near Weiser is located within the Rebecca Sand Hill RNA. These ACEC's support some of the largest and most extensive populations known. Due to the establishment of the ACEC's and the protection they afford, the long-term persistence of Aase's onion in the western half of its distribution now appears secure. The eastern half of its distribution remains vulnerable, however, especially in light of urban development patterns and the preponderance of private lands in the Boise Foothills. The BLM has also sponsored several Challenge cost-share projects to investigate and clarify the taxonomic disposition of Allium aaseae (McNeal 1993; Smith 1995), and has completed extensive field surveys to define its range and abundance. Other entities such as the Boise National Forest, Ada County, and Boise City have funded additional survey work in the Boise Foothills area (Moseley and Caicco 1989; Mancuso and Moseley 1991; Moseley et al. 1992).
The majority of Allium aaseae populations occur on private land, however, and subject to few conservation options. Conservation agreements involving private entities have not been pursued to any great degree. A proposed Conservation Agreement between the U. S. Fish and Wildlife Service and the City of Boise has been at a standstill for a prolonged period. When and if this Conservation Agreement is completed, and plans implemented, populations within Military Park Reserve (006, 025, 058, 059) and Camelsback Reserve (009) should be better protected than at present. One success involves Unimin Mining Corporation, which has been sponsoring propagation research for several years. Mixed ownership patterns and the preponderance of private lands throughout the range of Allium aaseae indicates cooperation, coordination, and innovation will be important for the species long-term viability. A lack of on-the-ground commitment from the private sector would be a serious drawback to the conservation of rare plants in the Boise Foothills. If this proves to be the case, populations on public land will be invaluable for the long-term persistence of Aase's onion and other rare plants found in the Boise Foothills.
Idaho: The Idaho Native Plant Society places Allium aaseae in the globally rare category of the state's rare plant list.
Global: Aase's onion is endemic to southwestern Idaho, occurring in the foothills around Boise and arcing northwest to near Emmett, an aerial distance of approximately 18 miles. In the Boise Foothills, the easternmost populations are known from the Hulls Gulch and lower Cottonwood Creek areas, while the Freezeout Hill vicinity near Emmett contains the westernmost foothill populations. Disjunct populations have recently been confirmed from near the towns of Payette and Weiser, northwest of the species' main range. Populations previously reported from the Danskin Mountains, east of Boise are really Allium simillimum. Populations are located in Ada, Boise, Gem, Payette, and Washington counties.
Idaho: See Global Distribution comments.
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| Allium aaseae habitat |
Elevation (Global): 2700 to 5100 feet
Elevation (Idaho): 2700 to 5100 feet
Global: Aase's onion is restricted to a narrow range of habitat conditions. It occurs on open, relatively barren, xeric, gentle to very steep, sandy slopes, generally with a southerly aspect, but ranging from east to west. It is usually associated with relatively sparsely vegetated bitterbrush (Purshia tridentata) or bitterbrush/sagebrush (Artemisia tridentata) communities. One or several bunchgrasses such as red threeawn (Aristida longiseta), bluebunch wheatgrass (Agropyron spicatum), squirreltail (Sitanion hystrix), needle-and-thread (Stipa comata), Sandberg's bluegrass (Poa sandbergii), Indian ricegrass (Oryzopsis hymenoides) and sand dropseed (Sporobolus cryptandrus) are often closely associated. Aase's onion sites are often bordered by Artemisia tridentata ssp. wyomingensis or ssp. tridentata/bunchgrass–dominated communities. Commonly associated species include Eriophyllum lanatum, Balsamorhiza sagittata, Achillea millefolium, Phacelia heterophylla, and Eriogonum ovalifolium. A number of exotic species may be abundant, especially Bromus tectorum, Erodium cicutarium and Taeniatherum caput-medusae.
Allium aaseae populations in the Boise Foothills often occur in close proximity to Astragalus mulfordiae, and/or Lepidium papilliferum, two other rare, regional endemic plants. These three rare species largely share the same conservation concerns and problems. On a local scale, Allium aaseae can be very common. At some sites it is one of the dominant forbs in early spring. When considering its sagebrush-bitterbrush/steppe and foothill grassland habitats rangewide, however, it is a minor constituent.
Most populations are restricted to the alluvial soils of the Glenns Ferry Formation. This sandy substrate is of granitic origin and typically coarse textured, well-drained and relatively deep (Packard 1979; Prentice 1988). In the Boise Foothills, all populations occur on one of three sand-dominated geologic units-Pierce Gulch Formation Sand, Terteling Springs Formation Sand and Sandstone, and Terteling Springs Formation Sandy Sediments (Beck 1988). A large majority of Boise Foothill populations occur on three soil mapping units of Beck (1988): Quincy-Lankbush complex, Payette-Quincy complex, and Haw-Lankbush complex. Rarely, populations or portions of populations occur on other soil types, namely, Lankbush-Brent sand loam, Ada gravelly sand, and Searless-Rock outcrop complex. All known populations except for the two in Cartwright Canyon occur between 2700 - 4300 feet elevation, with the great majority below 3700 feet. Cartwright Canyon populations occur at 4950 and 5100 feet, and possibly indicate that soil characteristics such as texture are more important than elevation in determining the distribution of Allium aaseae (McNeal 1993).
Idaho: See Global Habitat comments.
Global: There is little quantitative data regarding the effects of herbivores, disease, competition, hybridization or allelopathy on population viability. No native plant species appear to substantially compete with Allium aaseae for moisture, and only red three-awn seems to compete for space (Prentice 1988). Two exotic winter annuals, cheatgrass and storksbill, apparently are important interspecific competitors. Vigor of Allium aaseae populations can be reduced where these weeds are prolific (Prentice 1988). Livestock grazing on Allium aaseae is minimal, although indirect effects, such as habitat degradation and trampling are more serious. Deer have been observed feeding on Allium aaseae in early spring and chukars are known to eat bulbs later in the spring. The most serious insect pest seems to be an unknown seed predator that bores into and devours inner portions of the seed (Prentice 1988). A rust is common on populations in the Woods Gulch area, and maybe other places as well. The deep-seated bulb of Allium aaseae would survive wildfires. Hybridization and introgression are likely occurring between Allium aaseae and the more widespread A. simillimum (Smith 1995).
Idaho: See Global Ecology comments.
Global: Allium aaseae reproduces from both seed and bulb division. Seed viability from different sites is variable, as is often the case in wild pant populations. The mean viability from four sites studied by Prentice (1988) was 55%. The number of seeds per pound is estimated to be 622,000 (Prentice 1988). Seed production is also variable from year-to-year (Prentice 1989). The pollination biology of Allium aaseae is unknown, although it has been reported to have no specific pollinators, and flowers visited by many types of insects (Bolin and Rosentreter 1986). This species flowers early in the season and likely makes use of any insect taxa active at this time of year. Seed dispersal mechanisms are unknown, but probably at least partly relies on the dried, detached umbel being blown around (Packard 1979).
Idaho: See Global Reproduction comments.
Phenology (Idaho): Seed germination typically begins in late winter when snow melts, soil moisture increases and temperatures are cool. In late February, seedling roots emerge and the cotyledon, with the seed coat at the end of the single leaf, pushes up through the soil. The bulb begins to develop when the leaf is 1 - 2 cm long. It starts as a tiny bump on the root about 1.5 cm below the surface. The plant continues to grow until the leaf withers due to increasing moisture and temperature stresses. First- and second-year plants produce only one leaf, the bulb does not divide, nor produce flowers. Presence of a second leaf indicates an older individual, and appearance of a third and eventually a fourth leaf indicates the bulb is dividing (Prentice 1988; 1989).
Established plants (from bulbs) begin root growth and emerge in late winter (generally late February - early March, but in some years as early as late January). In mature bulbs, flower buds develop at ground level between the two leaves soon after emergence, and Allium aaseae is one of the first native plants to flower in the Boise Foothills (Prentice 1989). Anthesis varies from plant to plant, but seems to peak in early spring in most years. Higher elevation populations flower later than lower sites. In dry years, flowers are quite ephemeral, and may scarcely last a week, while during wet springs flowering plants can be found into May. Seeds can usually be found by late March (Packard 1979). Plants are dormant much of the year, generally from about May until February.
Global: Populations occur on private, City of Boise, Ada County, State Department of Lands, and BLM lands, often in mixed ownership. For the 68 known Allium aaseae occurrences, 31 (46%) occur solely on private land, while five (7%) are restricted to City of Boise land, two (3%) to Ada County land, four (6%) to State land, and two (3%) to BLM land. Where land ownership is mixed, 14 occurrences (20%) are on private and BLM land, one (2%) is on BLM and State land, one (2%) is on BLM and Ada County land, and 8 (12%) are on private and some other entity or mix of entities (Ada County, State, BLM). Lands under private ownership dominate for Allium aaseae. Fifty-two (76%) of the known occurrences are at least partly on private land. The other principal land owner is the BLM, with 18 (26%) occurrences at least partially on land they administer.
In 1993, an amendment to the BLM's Cascade Resource Management Plan designated six Areas of Critical Environmental Concern (ACEC's) to specifically protect populations of Allium aaseae. The six ACEC's are Cartwright Canyon (037), Sand-capped Knob (049), Sand Hollow (034), Willow Creek (039), Woods Gulch (053) and Hulls Gulch. Recent taxonomic analysis has revealed the Hulls Gulch population to be the closely related taxon Allium simillimum, or a more likely, a hybrid. One disjunct population (018) of Allium aaseae near Weiser is located within the BLM's Rebecca Sand Hills Research Natural Area (RNA), and is therefore protected. For the five BLM ACEC's containing Allium aaseae, a monitoring plan has been prepared in consultation with the USFWS. Monitoring plots have been established and baseline trend and habitat data was collected in 1991, 1992, 1994 (Bureau of Land Management 1992).
Populations on City of Boise, and BLM land occur within areas already, or scheduled to be at least partly dedicated to the conservation of Allium aaseae. A Conservation Agreement for the conservation of Allium aaseae on Boise City lands is pending between the U. S. Fish and Wildlife Service and Boise City Park Department. This Agreement will potentially affect populations in Military Reserve Park (006, 025, 058, 059), Camelsback Reserve (009) and lower Hulls Gulch (011).
A Cooperative Agreement between the BLM and Unimin Corporation regarding the propagation of Allium aaseae was signed in 1987. Portions of the Unimin Corporation sand mining operations near Emmett support an extensive population of Allium aaseae (015). Propagation and related research is ongoing, even though Unimin now owns the land (it was patented in 1993).
In 1992, the Idaho Conservation Data Center completed a contract with Boise City Planning and Zoning to conduct rare plant inventories in the Boise Foothills. This study provided rare plant population and habitat information, one of several information gaps hindering formulation of a comprehensive foothill planning document. Presently, the Boise Foothills Plan is under review by the Planning and Zoning Commissions of both Boise City and Ada County. The adoption of a final Foothills Plan has been delayed several times and continues to be controversial. It is unclear when this Plan will be finalized. At this stage, it is also unclear how strong the Plan will be regarding rare plant conservation, but early indications are not encouraging in this regard (a policy statement supporting the protection of rare plant populations is expected, but there may be little concerning techniques/options of how to do so). During the several years interim between completion of the inventory and expected Plan adoption, there apparently has been no consistent policy for the protection of rare plants such as Allium aaseae in foothill areas scheduled for development.
There is also a MOU currently being drafted addressing the implementation of consistent and uniform regulations and management for the foothills area. This MOU will likely include City of Boise, Ada County, Idaho Department of Lands, State Department of Agriculture, Agricultural Research Service, U.S. Forest Service, BLM, Idaho Department of Fish and Game and other entities. What emphasis or role this will have concerning rare plant conservation is unknown. Due to the nature of the 1872 General Mining Law, management options and regulations are limited regarding sites on public land supporting mining claims. The commercial quality of Glenns Ferry Formation sand deposits, especially around Emmett, will likely ensure sand mining will continue within the range of Allium aaseae. A draft Habitat Conservation Assessment (Mancuso 1995) and Conservation Strategy (Mancuso 1995) have been prepared for Allium aaseae as part of the Idaho Conservation Effort.
Idaho: See Global Management comments.
General Comments (Idaho):
1986 to Present: The BLM and Unimin Mining Corporation have conducted field surveys associated with ongoing propagation and life history studies in the Emmett area (Prentice 1988; 1989).
1986 to Present: The BLM has conducted intensive surveys for Allium aaseae throughout its range, predominantly on BLM land. The most thorough surveys took place in 1987 and 1988. Periodic, less intensive field surveys continue, often as part of project clearances and related work.
1987: The Conservation Data Center, under contract from Ada County Solid Waste Management, determined the status and distribution of Allium aaseae on county land in Seaman Gulch, an area used as a county landfill (Moseley and Caicco 1989).
1991: The Conservation Data Center, Idaho Native Plant Society, and Friends of Military Reserve mapped the distribution and abundance of Allium aaseae in Military Reserve Park, Boise.
1991: The Idaho Native Plant Society, Wetlands Coalition, and Golden Eagle Audubon mapped the distribution and abundance of Allium aaseae in lower Hulls Gulch, Boise.
1991: The Idaho Native Plant Society and Conservation Data Center mapped the distribution and abundance of Allium aaseae in Camelsback Reserve Park, Boise.
1991: The Conservation Data Center, as a cooperative Challenge cost-share project with the Boise National Forest, conducted a status survey for Allium aaseae on the Boise National Forest (Mancuso and Moseley 1991).
1992: The Conservation Data Center, under contract from Boise City Planning and Zoning Department, conducted field surveys for Allium aaseae throughout much of the Boise Foothills as one part of a rare plant and riparian inventory for the area (Moseley et al. 1992).
1993: Dr. Dale McNeal (University of the Pacific), in a cooperative Challenge cost-share project between the Conservation Data Center and BLM (and involving U. S. Fish and Wildlife Section 6 funding), conducted field work in association with his taxonomic investigation of Allium aaseae (McNeal 1993).
1994: Dr. Jim Smith (Boise State University) revisited a number of sites to support his genetic study of the Allium aaseae. This was another cooperative Challenge cost-share project with the BLM (Smith 1995).
1995: The Conservation Data Center conducted field investigations in the disjunct, Weiser and Payette portions of the species range. This was also part of a cooperative Challenge cost-share project with the BLM.
Inventory Needs (Idaho): There have been many field surveys for Allium aaseae over the years, and its limited distribution is well documented. Portions of the Boise foothills have never been thoroughly searched due to the large amount of private land in some areas. There is also additional unsurveyed habitat in the Weiser/Payette area, especially across the Snake River in Oregon.
Bolin, R., and R. Rosentreter. 1986. The autecology of Allium aaseae. Prepared for Unimin Corporation, New Canaan, CT. 12 pp.
Hitchcock, C. L., A. Cronquist, M. Ownbey, and J. W. Thompson. 1969. Vascular plants of the Pacific Northwest. Part I: Vascular cryptogams, gymnosperms and monocotyledons. University of Washington Press, Seattle. 914 pp.
Holsinger, K. E. 1978. The role of selenium in the evolution of Allium aaseae Ownbey. Unpublished paper. 21 pp.
Mancuso, M. 1995. Draft conservation strategy for Allium aaseae Ownbey (Aase's onion). Conservation Data Center, Idaho Department of Fish and Game, Boise. 8 pp. plus appendices.
Mancuso, M. 1995. Draft habitat conservation assessment for Allium aaseae Ownbey (Aase's onion). Conservation Data Center, Idaho Department of Fish and Game, Boise. 19 pp. plus appendices.
Mancuso, M., and R. K. Moseley. 1991. Field investigation of Allium aaseae (Aase's onion), on the Boise National Forest. Idaho Department of Fish and Game, Conservation Data Center, Boise. 14 pp. plus appendices.
McNeal, D. W. 1993. Taxonomy of Allium aaseae-Allium simillimum in Idaho. Unpublished report prepared for the Idaho Department of Fish and Game, Conservation Data Center. 10 pp.
Moseley, R. K., M. Mancuso, and J. Hilty. 1992. Rare plant and riparian vegetation inventory of the Boise Foothills, Ada County, Idaho. Idaho Department of Fish and Game, Conservation Data Center, Boise. 20 pp. plus appendices.
Moseley, R. K., and S. L. Caicco. 1989. Status and distribution of Aase's onion (Allium aaseae), a federal candidate species, on Ada County lands in Seaman Gulch. Prepared for Ada County Solid Waste Management. Idaho Department of Fish and Game, Conservation Data Center, Boise. 6 pp. plus appendices.
Packard, P. L. 1979. Status report for Allium aaseae. The College of Idaho, Caldwell. 11 pp.
Prentice, C. 1988. Progress report: a study of the life cycle of Allium aaseae Ownbey, Aase's onion. Coop agreement between Unimin Corporation and USDI Bureau of Land Management. 34 pp.
Prentice, C. 1989. 1989 Progress Report: A study of the life cycle of Allium aaseae Ownbey, Aase's onion. Cooperative Agreement between Unimin Corporation and USDI Bureau of Land Management, Boise, ID. 28 pp.
Smith, J. F. 1995. The genetic diversity of the rare Idaho endemic Allium aaseae Ownbey (Alliaceae) and potential introgression with A. simillimum Henderson. Cooperative Challenge Cost Share Project, Bureau of Land Management, Boise District Office, and Boise State University, Biology Department. 186 pp.
Author: M. Mancuso
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